Unit 2: Taxonomic Hierarchy & Botanical Nomenclature
Principles and Rules of ICN
The International Code of Nomenclature for algae, fungi, and plants (ICN) is the set of rules and recommendations dealing with the formal botanical names that are given to plants. Its intent is that each taxonomic group of plants has only one correct name that is accepted worldwide.
Core Principles of ICN
- Independence: Botanical nomenclature is independent of zoological and bacteriological nomenclature.
- Typification: The application of names of taxonomic groups is determined by means of nomenclatural types.
- Priority: The nomenclature of a taxonomic group is based upon priority of publication.
- Uniqueness: Each taxonomic group with a particular circumscription, position, and rank can bear only one correct name.
- Scientific Names: Scientific names of taxonomic groups are treated as Latin regardless of their derivation.
Ranks and Names
Taxonomic groups are treated in a hierarchical system of ranks. The primary ranks are Kingdom, Division, Class, Order, Family, Genus, and Species.
Suffixes for Specific Ranks
Typification and Author Citation
Typification
Typification is the process of designating a specific specimen (the type) to which the name of a taxon is permanently attached.
- Holotype: The single specimen designated by the author at the time of publication.
- Isotype: A duplicate specimen of the holotype, collected at the same time and place.
- Lectotype: A specimen chosen from the original material to serve as the type when no holotype was designated or is lost.
- Neotype: A specimen chosen to serve as the type when all original material is missing.
Author Citation
For a name to be complete and accurate, it should be followed by the name of the author who first validly published it. Example: Solanum tuberosum L. (where 'L.' stands for Linnaeus).
Valid Publication and Principle of Priority
Valid Publication
To be validly published, a name must be:
- Effectively published (distributed in printed matter to the public/institutions).
- Accompanied by a description or diagnosis of the taxon.
- Formulated according to the rules of the ICN.
Principle of Priority
If a taxon has been given multiple names, the oldest validly published name is the correct one. This principle starts from May 1, 1753 (Linnaeus's Species Plantarum). Limitations: Priority does not apply to ranks above family.
Naming of Hybrids
Hybrids are named using an 'x' sign.
- Interspecific: Fragaria x ananassa (Hybrid between two species).
- Intergeneric: x Triticosecale (Hybrid between Triticum and Secale).
Taxonomic Documentation
Documentation is essential for storing and retrieving information about plant diversity.
- Flora: A book describing the plants of a specific geographical area (e.g., Flora of British India).
- Monograph: A comprehensive study of a single taxonomic group (usually a genus or family) across its entire global range.
- Journals: Periodical publications where new taxa are described and taxonomic research is shared.
Taxonomic Keys
Keys are artificial devices used for the rapid identification of plants.
- Single Access (Dichotomous) Keys: Provide two contrasting choices (couplets) at each step.
- Indented Key: Each pair of choices is indented under the previous pair.
- Bracketeded Key: The two choices of a couplet are placed on adjacent lines.
- Multi-access (Polyclave) Keys: Allow the user to choose characters in any order, often used in computer-based identification systems.
Exam Tips
- Typification: Be very clear on the differences between Holotype, Isotype, and Lectotype. This is a common objective/short note topic.
- Priority: Remember the starting date (1753) for botanical priority.
- Keys: Be prepared to write a simple dichotomous key for a few common plants.
Common Mistakes
- Confusing **Effective Publication** (distribution) with **Valid Publication** (meeting all ICN requirements).
- Forgetting the 'x' when writing the scientific name of a hybrid.
Frequently Asked Questions
Q: What is a diagnosis in nomenclature?
A: A diagnosis is a statement that sets the taxon apart from its most closely related relatives by highlighting its unique features.
Q: Can a name be valid if it doesn't have a type?
A: No, the principle of typification requires that a name be associated with a physical type specimen for it to be validly applied.